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SGS

Shortgrass Steppe

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Sunset behind one of the still active windmills on the shortgrass steppe that provide water for livestock (1990s). Photo by Paul Stapp
SGS
Our SGS site encompasses a large portion of the Colorado Piedmont Section of the western Great Plains. The extent is defined as the boundaries of the Central Plains Experimental Range (CPER), which is managed by the Agricultural Research Service (ARS), and the Pawnee National Grassland (PNG), which is managed by the US Forest Service. Expansion into the PNG has allowed us to explore the biotic interactions of the SGS ecosystem across a range of climatic, geologic, topographic and land use conditions. The CPER has a single ownership and landuse (livestock grazing). The PNG is characterized by a mosaic of ownership and land use. Ownership includes federal, state or private and land use consists of livestock grazing or row-crops. There are NGO conservation groups that exert influence over the area, particularly on federal lands. This varied land use and diversity associated with land users and managers substantiates the importance of the sgs-lter to the area.

Climate

The climate of the SGS is typical of mid-continental semiarid regions in the temperate zone except for the strong influence of the Rocky Mountains approximately 60 km to the west. Mean monthly temperatures range from -4 to 22 degrees C seasonally and have a daily average max-min range of 17 degrees C. Annual precipitation at the CPER averaged 322 mm over the past 51 years and ranged between 107 and 588 mm. Approximately 70% of the mean annual precipitation occurs during the April to September growing season. ( For further information please see Figures Historical Precipitation, Temperature/Precipitation, and Summary Climate Information : SGS climate description and summary tables and charts)

Vegetation

The vegetation of the SGS is dominated by shortgrasses (64%), forbs (7%), succulents (21%), and half-shrubs (8%). The key species of these groups are Bouteloua gracilis and Buchloe dactyloides; Sphaeralcea coccinea; Opuntia polyacantha; and Chrysothamnus nauseosus, Gutierrezia sarothrae, and Eriogonum effusum, respectively. Average above ground net primary production is 125 g/m2 and ranges from 60 to 180 g/m2 depending on available soil water. Major differences in vegetation structure occur in saltgrass meadows dominated by Distichlis stricta and Sporobolus asper, and on floodplains where the shrub Atriplex canescens is an important component.
Facilities

The main LTER headquarters building (214 m2) has offices, laboratories, a dining/meeting room, and a kitchen. Adjacent to the headquarters is a storage/sample processing building (134 m2) with facilities for washing and drying samples. The dormitory has six rooms; five capable of double occupancy and one with four beds. In addition there are large-animal handling and holding pens and a residence for the site manager. The Shortgrass Steppe LTER Field Station was built to support research as part of the US/IBP Grassland Biome project in the early 1970s.

LTER headquarters building

The LTER Field Station is located 1 km east of State Highway 85 on Weld County Road 114. The station is 8 km north of Nunn, approximately halfway between Greeley, Colorado and Cheyenne, Wyoming, on the Central Plains Experimental Range (CPER). The CPER is a 6200-ha research area maintained by the USDA Agricultural Research Service for applied rangelands research. Field headquarters for the ARS/CPER are located 8 km NE of the LTER Field Station, on County Road 37.

For information on using the LTER Field Station and its facilities, contact Mark Lindquist , LTER Site Manager.

Short history: 
The SGS LTER project represents the continuing development of a research tradition that began with the US/IBP Grassland Biome project in the late 1960's, the time at which ecosystem science was formally recognized as a sub-discipline in ecology. Research at the CPER over the past 20 years has had an important interactive relationship with the development of ecosystem science. The Grassland Biome project focused on the issue of productivity of natural ecosystems. Grasslands were conceptualized as homogeneous entities, appropriately described by an average square meter. The transition from the IBP project in the early 1970's to the LTER project in the early 1980's involved a change in thinking about the importance of spatial variability.
History: 

Our involvement in the LTER program (LTER I 1982-1986) began with spatially explicit ideas and questions about the importance of landscape structure, particularly the classic soil catena model, in the long-term development and maintenance of shortgrass steppe ecosystems. In the second phase of the project (LTER II 1987-1990) we expanded our concept of long-term processes to include the origin and persistence of spatial patterns at a range of spatial scales (Fig 1). This work included substantial questioning of the generality of the catena model at the CPER and in the shortgrass steppe region. Our work for LTER III builds upon LTER I and II and expands the depth of our investigations into interactions between spatial and temporal patterns in ecosystem structure and function.

LTER IV will expand our understanding of SGS ecosystem structure and function, by continuing a substantial suite of long-term experiments and initiating new long-term monitoting, long- and short-term experiments, and simulation analyses. Our work will focus on the key biotic responses and feedbacks as influenced by humans, natural disturbances, physiography, and climate. Our work includes: 1) new studies of invasive species, the prairie dog and its biology, and cactus; 2) new studies of control by landscapes over biogeochemical cycling, new studies of the N balance of SGS ecosystems, and 3) new studies of carbon balance over landscape to regional scales, and the interactions between landuse and regional climate. We are also expanding experiments associated with our past work in paleoecology and pedology, plant community dynamics, and biogeochemistry. We plan to continue our 60 long-term experiments, and to initiate 14 new long-term experiments and 10 new short-term, synthesis and cross site experiments over the next six years.

Short research topics: 
The shortgrass steppe (SGS) LTER has been in operation since 1982. The key questions that guide our work are: 1. What factors regulate SGS ecological structure and function over space and time? 2. How do the factors that regulate ecological structure and function, and the coupling of biotic and abiotic components, vary spatially and temporally within the SGS? 3. What are the biotic and abiotic thresholds that determine the vulnerability of the SGS to changes in the factors influencing ecological structure and function?

Our conceptual framework asserts that SGS ecological structure and function are governed by climate, natural disturbance, physiography, human use, and biotic interactions. SGS-LTER work is divided into three topic areas, 1) Population Dynamics (plants and heterotrophs), 2) Biogeochemical Dynamics, and 3) Land-Atmosphere Interactions. Disturbances are of such importance that they are embedded in each of our topic areas. In addition to continuing our long-term experiments that address our key questions, we are initiating new studies that primarily focus on Question 3. The discussion below focuses mostly on this new work.

Population Dynamics: Plant Dynamics and Ecosystem Interactions: One grass species, Bouteloua gracilis, contributes most to biomass and net primary production (NPP), and has been the focus of most of our past work. We will initiate long-term studies of Opuntia polyacantha (prickly-pear cactus) a species that makes up a small proportion of NPP; its spiny growth form affords protection to other species and therefore may have a large effect under cattle grazing. Even though the SGS has proven quite resistant to invasive plants, the introduction of species that can successfully invade the SGS is inevitable. We propose to study the potential for invasion by plants with different life history attributes, and to survey extensively for new invasions.

Faunal Dynamics: The dynamics of small mammals simultaneously reflect and affect both the structure and function of the SGS ecosystem. We will continue our long-term studies tracking small mammal populations, their resources, and predators. New research will emphasize the dynamics of the black-tailed prairie dog (Cynomys ludovicianus) and its effects on biological diversity and ecosystem function. Prairie dogs were abundant in the past, but have been severely affected by poisoning, shooting, habitat destruction, and, most significantly, by an introduced bacterium that causes plague (Yersinia pestis). The prairie dog is now a candidate for listing under the Endangered Species Act. We will examine prairie dog dynamics as affected by plague.

Biogeochemical Dynamics: We conceive of biogeochemical processes as an integrated function of biotic components and those processes responsible for carbon and nutrient fluxes: primary production, decomposition, weathering, mineralization, and trace gas flux. We will continue to study variations in biogeochemistry as a function of climatic controls, and will initiate new studies that evaluate geologic (textural) and topographic controls, and balancing the N budget.

Land-Atmosphere: The processes with which we are primarily concerned are water, energy, and gaseous fluxes that represent important interactions between the ecosystem and the atmosphere. One of the most important alterations that has occurred and continues to occur on the SGS LTER is land use change. We will evaluate the interactions of human land use, biotic responses, and atmospheric processes at scales from local to regional.

Synthesis: The SGS has a major investment in simulation analysis, and each of our topic areas has simulation components. We will continue to synthesize long-term data both within and across research areas. Our new synthesis volume will be published during this next funding cycle.

Other Activities: Data management supports our research and contributes to ecoinformatics. We will continue to build our interactions with federal land management agencies, NGO’s, and the interested public. Through the use of supplemental funding, the SGS-LTER will focus on K-12 and community outreach through student mentoring, curriculum development, teacher professional development, community partnerships, and education research.

Dept. of Biology
College of Natural Sciences
Colorado State University
Fort Collins
CO
80523-1499
USA
970-491-2366
970-491-2156
Principal investigator name: 
John Moore
Grad rep name: 
Karie Cherwin
Climate rep name: 
William Lauenroth
Information manager name: 
Nicole Kaplan
Education contact name: 
John Moore
Social Science Rep: 
Catherine Keske
Primary contact name: 
Sallie Sprague
Shortgrass Steppe
elevation comment: 
Data Source: Collins/Waide. class data. 2008. not published yet.
latitude comment: 
Data Source: LTER Site Characteristics Database. http://www.lternet.edu/sites/sgs
Longitude_comment: 
Data Source: LTER Site Characteristics Database. http://www.lternet.edu/sites/sgs
ecosystem comment: 
Data Source: GreenLand, D., G. G. Goodin., R. C., Smith. 2003. An Introduction to Climate Variability and Ecosystem Response. p8. In Climate Variability and Ecosystem Response at Long-Term Ecological Research Sites. Oxford University Press

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